Papers

Ochi & Araya

Studies on the Coprophagous Scarab Beetles from East Asia. II (Coleoptera, Scarabaeidae)

1992 Giornale Italiano di Entomologia

Studies on th Coprophagous Scarab Beetles from East Asia. III (Coleoptera, Scarabaeidae)

1992 Giornale Italiano di Entomologia

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NOTE: Where applicable, species list above only shows those found in the Philippines.